FinalLeafNumberSet
The FinalLeafNumberSet model stores final leaf number (FLN) observations or estimates for a genotype under controlled environmental conditions. These observations help quantify the effects of vernalisation and photoperiod treatments on leaf development, supporting genotype-specific modelling of phenology in APSIM NG.
Overview
FinalLeafNumberSet defines the final leaf number (FLN) under four treatments in the controlled environments (Bloomfield et al. 2023):
- LV: fully vernalised long photoperiod
- SV: fully vernalised short photoperiod
- SN: un-vernalised short photoperiod
- LN: un-vernalised long photoperiod
This model is typically used within the CAMP framework to parameterise genotype responses.
Inputs
This model does not accept dynamic inputs during simulation, but rather defines fixed parameters:
- MinLN: Base final leaf number under full vernalisation and long photoperiod.
- PpLN: Increase in FLN when grown under short photoperiod (e.g. 8h) after full vernalisation.
- VrnLN: Increase in FLN due to lack of vernalisation when grown under short photoperiod.
- VxPLN: Modification to FLN due to photoperiod interaction under unvernalised conditions.
These values are typically derived from controlled environment experiments.
Methodology
The model exposes computed FLN values based on defined treatment combinations:
- LV (Long photoperiod, fully vernalised):
\[ \text{LV} = \text{MinLN} \]
- SV (Short photoperiod, fully vernalised):
\[ \text{SV} = \text{MinLN} + \text{PpLN} \]
- SN (Short photoperiod, un-vernalised):
\[ \text{SN} = \text{MinLN} + \text{PpLN} + \text{VrnLN} \]
- LN (Long photoperiod, un-vernalised):
\[ \text{LN} = \text{MinLN} + \text{VrnLN} + \text{VxPLN} \]
These equations represent the phenotypic plasticity of FLN in response to vernalisation and photoperiod interactions.
Reversely, the model parameters can be derived from the final leaf number (FLN) values observed in the controlled environment experiments:
- MinLN:
\[ \text{MinLN} = \text{LV} \]
- PpLN:
\[ \text{PpLN} = \text{SV} - \text{LV} \]
- VrnLN:
\[ \text{VrnLN} = \text{SN} - \text{SV} \]
- VxPLN:
\[ \text{VxPLN} = \text{LN} - \text{LV} - \left(\text{SN} - \text{SV}\right) \]
Events
Events Listened For
No events are listened by this function.
Events Raised to
No events are raised by this function.
Properties
Configurable and Reportable Properties
| Property | Type | Description |
|---|---|---|
| MinLN | double | Final Leaf Number under full vernalisation and long photoperiod (>16h) |
| PpLN | double | Increase in FLN under short photoperiod (<8h) following full vernalisation |
| VrnLN | double | Increase in FLN when un-vernalised and grown under short photoperiod |
| VxPLN | double | Change in FLN from photoperiod effect under un-vernalised, long-day growth |
Read-Only Reportable Properties
| Property | Type | Description |
|---|---|---|
| LV | double | FLN under full vernalisation and long photoperiod (>16h) |
| SV | double | FLN under full vernalisation and short photoperiod (<8h) |
| SN | double | FLN when un-vernalised under short photoperiod |
| LN | double | FLN when un-vernalised under long photoperiod |
User Interface
FinalLeafNumberSet can be added as a child of a CAMP node in the model tree. Right-click the CAMP node, select “Add Model…”, and search for FinalLeafNumberSet in the Filter Box.
Practical Example
Below is a summary table for two New Zealand cultivars (i.e. BattenWinter and BattenSpring), showing their parameter values and calculated FLN for each treatment:
| Parameter / FLN | BattenWinter | BattenSpring |
|---|---|---|
| MinLN | 8.16 | 8.25 |
| PpLN | 3.61 | 7.55 |
| VrnLN | 3.10 | 0.19 |
| VxPLN | 6.99 | -0.03 |
| LV | 8.16 | 8.25 |
| SV | 11.77 | 15.80 |
| SN | 14.87 | 15.99 |
| LN | 18.25 | 8.41 |
This figure illustrates the final leaf number (FLN) for the BattenWinter and BattenSpring cultivars under different treatments, including the contributions of each parameter to the final leaf number.
See Also
- Source code: FinalLeafNumberSet.cs on GitHub
References
Bloomfield, Maxwell T., Corinne Celestina, James R. Hunt, Neil Huth, Bangyou Zheng, Hamish Brown, Zhigan Zhao, et al. 2023. “Vernalisation and Photoperiod Responses of Diverse Wheat Genotypes.” Crop and Pasture Science, March. https://doi.org/10.1071/CP22213.